The very first thing that Bezalel constructed was the Ark of the Covenant, contrary to Moses' order, first to erect the Tabernacle and then to supply its separate furnishings. He succeeded in convincing Moses that it was the proper thing to begin with the Ark, saying: "What is the purpose of this Tabernacle?" Moses: "That God may let His Shekinah rest therein, and so teach the Torah to His people Israel." Bezalel: "And where dost thou keep the Torah?" Moses: "As soon as the Tabernacle shall have been complete, we shall make the Ark for keeping the Torah." Bezalel: "O our teacher Moses, it does not become the dignity of the Torah that in the meanwhile it should lie around like this, let us rather first make the Ark, put the Torah into it, and then continue with the erection of the Tabernacle, for the Tabernacle exists only for the sake of the Torah." Moses saw the justice of this argument, and Bezalel began his work with the construction of the Ark. In this he followed the example of God, who created light before all the rest of the creation. So Bezalel first constructed the Ark that contains the Torah, the light that illuminates this world and the other world; and only then followed the rest.  The Ark consisted of three caskets, a gold one, the length of then spans and a fractional part; within this a wooden one, nine spans long, and within this wooden one, one of gold, eight spans long, so that within and without the wooden was overlaid with the golden caskets. The Ark contained the two tables of the Ten Commandments as well as the Ineffable Name, and all His other epithets. The Ark was an image of the celestial Throne, and was therefore the most essential part of the Tabernacle, so that even during the march it was spread over with a cloth wholly of blue, because this color is similar to the color of the celestial Throne. It was through the Ark, also, that all the miracles on the way through the desert had been wrought. Two sparks issued from the Cherubim that shaded the Ark, and these killed all the serpents and scorpions that crossed the path of the Israelites, and furthermore burned all thorns that threatened to injure the wanderers on their march through the desert. The smoke rising from these scorched thorns, moreover, rose straight as a column, and shed a fragrance that perfumed all the world, so that the nations exclaimed: "Who is this that cometh out of the wilderness like pillars of smoke, perfumed with myrrh and frankincense, with all powders of the merchant?" 
Apart from this Ark, which was kept in the Tabernacle, they had another ark, in which were contained the tables broken by Moses, which they carried with them whenever they went to war.  The Ark that Bezalel constructed was also used again in Solomon's Temple, for he retained the Ark used by Moses in the Tabernacle, even though all the other furnishings of the Temple were fashioned anew. It remained there up to the time of the destruction of the Temple by Nebuchadnezzar, when it was concealed under the pavement of the wood-house, that it might not fall into the hands of the enemy. This place remained a secret for all time. Once a priest, noticing about the wood-house that something lay hidden under it, called out to his colleagues, but was suddenly stricken dead before divulging the secret. 
On the Ark were the Cherubim with their faces of boys and their wings. Their number was two, corresponding to the two tables, and to the two sacred names of God, Adonai and Elohim, which characterized Him as benevolent and as powerful. The face of each Cherub measured one span, and the wings extended each ten spans, making twenty-two spans in all, corresponding to the twenty-two letters of the Hebrew alphabet.  It was "from between the two Cherubim" that God communed with Moses, for the Shekinah never wholly descended to earth any more than any mortal ever quite mounted into the heaven, even Moses and Elijah stood a slight distance from heaven; for, "The heaven, even the heavens, are the Lord's: but the earth hath He given to the children of men." Therefore God chose the Cherubim that were ten spans above the earth as the place where the Shekinah betook itself to commune with Moses.  The heads of the Cherubim were slightly turned back, like that of a scholar bidding his master farewell; but as a token of God's delight in His people Israel, the faces of the Cherubim, by a miracle, "looked one to another" whenever Israel were devoted to their Lord, yea, even clasped one another like a loving couple. During the festivals of the pilgrimage the priest used to raise the curtain from the Holy of Holies to show the pilgrims how much their God loved them as they could see in the embrace of the two Cherubim. 
A tow-fold miracle came to pass when the Cherubim were brought into the Temple by Solomon: the two staves that were attached to the Ark extended until they touched the curtain, so that two protuberances like a woman's breasts became visible at the back of it, and the wings of the Cherubim furthermore extended until they reached the ceiling of the Holy of Holies.